Hart, Lachlan J.McCurry, Matthew RFrese, MichaelPeachey, Thomas J.Brocks, Jochen2023-08-270311-5518http://hdl.handle.net/1885/296882A single tetrapod tooth has been recovered from the Upper Jurassic Talbragar Fossil Fish Bed of New South Wales, Australia. It is the first evidence of a tetrapod to have been found at this locality in over 130 years of excavation. The tooth is likely from a temnospondyl amphibian. Herein, we document the discovery, discuss the potential explanations as to why tetrapod remains are so scarce from this locality and provide hypotheses as to how this tooth came to be preserved.We would like to thank Levi Beeching and Michael Turner (CTLab, ANU) for technical assistance, Tim Senden (ANU) for stimulating discussions and access to the scanning facilities, John Long for assistance in taxonomic identification, Nigel McGrath for allowing access to his property on which this fossil was found, and Patrick Smith (AM) for helpful discussions on the stratigraphy and lithology and for providing the locality map in Fig. 1. We thank CSIRO's Australian National Insect Collection for providing free access to imaging equipment. We appreciate the comments from the Chief Editor of Alcheringa, Benjamin Kear, as well as our reviewer Bryan Gee (University of Washington) and an anonymous referee, who improved the quality of the manuscript. This research was supported by an Australian Government Research Training Program (RTP) Scholarship to LJHapplication/pdfen-AU© 2021 The authorsTalbragarJurassicAustraliaTemnospondylitoothmicroCTplicidentine202110.1080/03115518.2021.19836512022-07-24